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watershed is also used in the sense of a boundary, isn't it?
Yes, there is certainly a more metaphorical usage for "watershed" too, meaning a moment or thing that divides what is being discussed into two distinct parts. "X was a watershed moment in the history of Y", for example. I think the author intends to use "ecotone" to represent a more gradual transition period -- a subtle filtering rather than a dramatic change.
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